Home / Prof. Dr. Katerina Harvati-Papatheodorou


Prof. Harvati is a paleoanthropologist specializing in Neanderthal evolution, modern human origins and the application of 3-D geometric morphometric and virtual anthropology methods to paleoanthropology.

Her broader research interests include primate and human evolution; evolutionary theory; evolution of primate and human life-history; the relationship of morphological variability to population history and the environment; and Paleolithic archaeology. She has conducted fieldwork in Europe and Africa, and recently directed paleoanthropological fieldwork in Greece and Tanzania.

 Her work has been published in Nature, Science, the Proceedings of the National Academy of Sciences, Evolutionary Anthropology, Journal of Human Evolution, American Journal of Physical Anthropology and other peer-reviewed international journals. She is the editor of two volumes in the Vertebrate Paleobiology and Paleoanthropology series of Springer Verlag: on Neanderthal paleobiology and on the Paleoanthropology of the Balkans.

 Her research was named one of the top 10 scientific discoveries of the year 2007 by TIME magazine for demonstrating the African origin of all modern humans (link to the article). In 2010 she was elected Fellow of the American Association for the Advancement of Science for her contributions to Paleoanthropology (link to the site). She was awarded the prestigious Research Award of the state of Baden-Württemberg in 2014 (basic research).

Prof. Harvati is the recipient of two ERC grants (ERC Starting Grant ‘Paleoanthropology at the Gates of Europe’, 2011; ERC Consolidator Grant ‘Human Evolution at the Crossroads’, 2016). In addition to her ongoing Consolidator Grant (CROSSROADS) she directs the DFG Centre for Advanced Studies ‘Words, Bones, Genes, Tools: Tracking linguistic, cultural and biological trajectories of the human past’ together with Prof. Gerhard Jäger (Linguistics).

 Since 2009 Prof. Harvati also holds an adjunct Professorship of Anthropology at the City University of New York Graduate Center and is resource faculty for the New York Consortium of Evolutionary Primatology.

Academic Background
2009Habilitation, Paleoanthropology. Eberhard Karls Universität Tübingen.
2001Ph. D. in Anthropology, City University of New York (CUNY) Graduate School and New York Consortium in Evolutionary Primatology
1998M.A. in Anthropology, Hunter College, CUNY
1994B.A. in Anthropology (Summa Cum Laude), Columbia College, Columbia University
Academic Positions
2009 - presentProfessor, Director of Paleoanthropology. Eberhard Karls Universität Tübingen
2005 - presentAdjunct Associate Professor, Department of Anthropology, City University of New York Graduate School
2001 - 2004Senior Researcher, Max Planck Institute for Evolutionary Anthropology
2000Assistant Professor, New York University
1997Adjunct Lecturer, Lehman College, CUNY
1995 - 1997Adjunct Lecturer, Lehman College, CUNY
Adjunct Lecturer, Hunter College, CUNY
Awards and Fellowships
2014Landesforschungspreis Baden-Württemberg for basic research
2010Elected Fellow of the American Association for the Advancement of Science (link)
2009Hellenes abroad award - Woman of the year 2009, Europe
2007TIME Magazine top ten scientific discoveries of 2007 (link)
2006Presentation Prize, 150 Years of Neanderthal Conference
2000 - 2001City University of New York Dissertation Year Fellowship.
1998 - 2000American Museum of Natural History Doctoral Fellowship.
1997 - 1999Alexander S. Onassis Foundation Doctoral Fellowship.
1998E. A. Hooton Student Prize, American Association of Physical Anthropologists
1994 -1997Research Fellowship, City University of New York / NYCEP
1993Stigler Fund Award, Department of Anthropology, Columbia University.
1993ΦBK, Rabi Scholar’s Society, Columbia University.
Edited Volumes
9789402408737Harvati K, Roksandic, M. (Eds.) 2016. Paleoanthropology of the Balkans and Anatolia: Human Evolution and its Context. Vertebrate Paleobiology and Paleoanthropology Series, Springer: Dordrecht.
84b3711434Harvati, K and Harrison, T. 2006. Neanderthals Revisited: New Approaches and Perspectives. Springer: Dordrecht. ISBN: 978-1-4020-5120-3
About the book
Publications in international refereed journals and edited volumes
Karakostis F. A., Hotz G., Tourloukis V. , Harvati K. 2018. Evidence for precision grasping in Neandertal daily activities. Science Advances.
Karakostis A.F., Hotz G., Scherf H., Wahl J., Harvati K. In press. Occupational manual activity is reflected on the patterns among hand entheses. American Journal of Physical Anthropology
Schuenemann V.J., Peltzer A., Welte B., van Pelt P.W., Molak M., Wang C.C., Furtwängler A., Urban C., Reiter E., Nieselt K., Teßmann B., Francken M., Harvati K., Haak W., Schiffels S., Krause J. 2017. Ancient Egyptian mummy genomes suggest an increase of Sub-Saharan African ancestry in post-Roman periods. Nature Communications 8,15694
Singh N., Albert F.W., Plyusnina I., Trut L., Pӓӓbo S., Harvati K. 2017. Facial shape differences between rats selected for tame and aggressive behaviors. PLOS ONE 12(4): e0175043.
Jean-Jacques Hublin, Abdelouahed Ben-Ncer, Shara E. Bailey, Sarah E. Freidline, Simon Neubauer, Matthew M. Skinner, Inga Bergmann, Adeline Le Cabec, Stefano Benazzi, Katerina Harvati, Philipp Gunz: New fossils from Jebel Irhoud, Morocco and the pan-African origin of Homo sapiens. Nature 546, 289–292 (08 June 2017) doi:10.1038/nature22336
Reyes-Centeno H, Ghirotto S, and Harvati K. 2017. Genomic validation of the differential preservation of population history in modern human cranial anatomy. Am J Phys Anthropol 162(1):170-179.
Reyes-Centeno H., Harvati K., Jäger G. 2016. Tracking modern human population history from linguistic and cranial phenotype. Scientific Reports. DOI: 10.1038/srep36645
Reyes-Centeno H., Ghirotto S., Harvati K. 2016. Genomic validation of the differential preservation of population history in the human cranium. American Journal of Physical Anthropology. DOI: 10.1002/ajpa.23060
Harvati K. 2016. Paleoanthropology in Greece: recent findings and interpretations. In: K. Harvati and M. Roksandic, eds. Paleoanthropology of the Balkans and Anatolia: Human Evolution and its Context. Vertebrate Paleobiology and Paleoanthropology Series, pp. 3-14. Springer: Dordrecht.
Harvati K. and Roksandic M. 2016. The Human Fossil Record from Romania: Early Upper Paleolithic Mandibles and Neanderthal Admixture. In: K. Harvati and M. Roksandic, eds. Paleoanthropology of the Balkans and Anatolia: Human Evolution and its Context. Vertebrate Paleobiology and Paleoanthropology Series, pp.51-68. Springer: Dordrecht
Aytek A.I. and Harvati K. 2016. The human fossil record from Turkey. In: K. Harvati and M. Roksandic, eds. Paleoanthropology of the Balkans and Anatolia: Human Evolution and its Context. Vertebrate Paleobiology and Paleoanthropology Series, pp. 79-91. Springer: Dordrecht.
Rathmann H., Semerari G.S., Harvati K. 2016. Evidence for migration influx into the ancient Greek colony of Metaponto: A population genetics approach using dental nonmetric traits. International Journal of Osteoarchaeology. DOI: 10.1002/oa.2569
Noback M.L., Samo E., van Leeuwen C.H.A., Lynnerup N., Harvati K. 2016. Paranasal sinuses: a problematic proxy for climate adaptation in Neanderthals. Journal of Human Evolution 97, 176-179.
Fu Q., Posth C., Hajdinjak M., Petr M., Mallick S., Fernandes D., Furtwängler A., Haak W., Meyer M., Mittnik A., Nickel B., Peltzer A., Rohland N., Slon V., Lazaridis I., Lipson M., Schiffels S., Skoglund P., Gély B., Benazzi S., Gonzales Morales M.R., Peresani M., Straus L.G., Caramelli D., Lari M., Ronchitelli A., Valentin F., Constantin S., Coppola D., Grigorescu D., Neugebauer-Maresch C., Thevenet C., Wehrberger K., Crevecoeur I., Rougier H., Semal P., Cupillard C., Bocherens H., Mannino M., Conard N.J., Harvati K., Moiseyev V., Drucker D.G., Svoboda J., Pinhasi R., Kelso J., Patterson N., Krause J., Pääbo S., Reich D. 2016. The genetic history of Ice Age Europe. Nature 524, 200-2005 doi:10.1038/nature17993
Posth C., Renaud G., Mittnik A., Drucker D.G., Rougier H., Cupillard C., Valentin F., Thevenet C., Furtwängler A., Wißing C., Franken M., Malina M., Bolus M., Lari M., Gigli E., Capecchi G., Crevecoeur I., Beauval C., Flas D., Germonpré M., van der Plicht J., Cottiaux R., Gély B., Ronchitelli A., Wehrberger K., Grigourescu D., Svoboda J., Semal P., Caramelli D., Bocherens H., Harvati K., Conard N.J., Haak W., Powell A., Krause J. 2016. Pleistocene human mitochondrial genomes suggest late dispersal of non-Africans and a Late Glacial population turnover in Europe. Current Biology 26, 827-833.
Tourloukis V., Thompson N., Garefalakis C., Konidaris G., Karkanas P., Panagopoulou E., Harvati K. 2016. New Middle Paleolithic sites from the Mani peninsula, Southern Greece. Journal of Field Archaeology 41, 68-83.
Uhl A., Reyes-Centeno H., Grigorescu D., Kranioti E.F., Harvati K. 2016. Inner ear morphology of the Cioclovina early modern European calvaria from Romania. American Journal of Physical Anthropology 160, 62-70.
Bauer C.C., Bons P.D., Benazzi S., Harvati K. 2016. Technical note: Using elliptical best fits to characterize dental variation. American Journal of Physical Anthropology 159, 342-347
Scherf H., Wahl J., Hublin J.-J., Harvati K. 2016. Patterns of activity adaptation in humeral trabecular bone in Neolithic people and present day humans. American Journal of Physical Anthropology 159, 106-115.
Konidaris G.E., Tourloukis V., Kostopoulos D.S., Thompson N., Giusti D., Mihailidis D., Koufos G.D., Harvati K. 2015. Two new vertebrate localities from the Early Pleistocene of Mygdonia Basin (Macedonia, Greece): Preliminary results. Comptes Rendus Palevol 14, 353-362
Reyes-Centeno H., Hubbe M., Hanihara T., Stringer C., Harvati K. 2015. Testing Modern Human Out-of-Africa Dispersal Models and Implications for Modern Human Origins. Journal of Human Evolution 87, 95-106
Panagopoulou E., Tourloukis V., Thompson N., Athanassiou A., Tsartsidou G., Konidaris G.E., Giusti D., Karkanas P., Harvati K. 2015. Marathousa 1 – A new Middle Pleistocene archaeological site from Greece. Antiquity Project Gallery 89 (343).
Noback M., Harvati K. 2015. The contribution of diet to global human cranial variation. Journal of Human Evolution 80, 34-50.
Harvati, K., Bauer, C.C., Grine, F.E., Benazzi, S., Ackerman, R.R., van Niekerk, K.L., Henshilwood, C.S. 2015. A human deciduous molar from the Middle Stone Age (Howiesons Poort) of Klipdrift Shelter, South Africa, Journal of Human Evolution, 82, 190-196
Bauer, C.C., Harvati, K. 2015. A virtual reconstruction and comparative analysis of the KNM-ER 42700 cranium. Anthropologischer Anzeiger 72(2), 129-140.
Harvati K. 2015. Neanderthals and their contemporaries. In: W. Henke and I. Tattersall (eds.) Handbook of Paleoanthropology, 2nd Edition. Springer.
Noback M., Harvati K. 2015. Covariation in the human masticatory apparatus. Anatomical Record, 291, 68-84.
Rein T.R., Harvati K., Harrison T. 2015. Inferring the use of forelimb suspensory locomotion by extinct primate species via shape exploration. Journal of Human Evolution 78, 70-79.
Hubbe, M., Neves, W.A., Harvati, K. 2015. Cranial morphology of early South Americans: Implications for understanding human dispersion into the New World. In: Frachetti, Michael David; Spengler III, Robert N. (Eds). Mobility and Ancient Society in Asia and the Americas. Springer, NY: pp. 103-116
Harvati K., Toussaint M. 2014. 3-D geometric morphometric analysis of the Scladina Neandertal child’s mandible in a developmental context. In M. Toussaint & D. Bonjean (eds.) The Scladina I-4A Juvenile Neanderthal (Andenne, Belgium) Paleoanthropology and Context. Études et Recherches Archéologiques de l’Université de Liège, 134, pp. 215-222.
Fornai, C., Benazzi, S., Svoboda, J., Pap, I., Harvati, K., Weber, G. W. 2014. Enamel thickness variation of deciduous first and second upper molars in modern humans and Neandertals. Journal of Human Evolution 76, 83-91.
Slizewski A., Burger-Heinrich E., Francken M., Wahl J., Harvati K. 2014. Pilot study for reconstruction of soft tissues in Archaeology: Muscle cross-sectional area of the forearm estimated from cortical bone for a Neolithic sample. Anatomical Record 297,1103-1114.
Rein T.R., Harvati K. 2014. Geometric Morphometrics and Virtual Anthropology: Advances in human evolutionary studies. Anthropologischer Anzeiger 71(1-2), 41-55
20.coverReyes-Centeno H., Ghirotto S., Détroit F., Grimaud-Hervé D., Barbujani G, Harvati K. 2014. Genomic and Cranial Phenotype Data Support Multiple Modern Human Dispersals from Africa and a Southern Route into Asia. Proceedings of the National Academy of Sciences USA 111, 7248-7253
Slizewski, A., Shönau, E., Shaw, C., Harvati, K. 2013. Muscle Area Estimation from Cortical Bone. Anatomical Record 296, 1695-1707.
Freidline S.E., Gunz P., Harvati K., Hublin J.-J. 2013. Evaluating developmental shape changes in Homo antecessor subadult facial morphology. Journal of Human Evolution 65, 404-423.
Baab K., McNulty K., Harvati K. 2013. Homo floresiensis Contextualized: A Geometric Morphometric Comparative Analysis of Fossil and Pathological Human Samples. PLoS1 8(7): e69119.
Scherf H., Harvati K., Hublin J.-J. 2013. A comparison of proximal humeral cancellous bone of great apes and humans. Journal of Human Evolution 65, 29-38.
Osipov B., Harvati K., Nathena D., Spanakis K, Karantanas A., Kranioti E.F. 2013. Sexual Dimorphism of the Bony Labyrinth: A New Age Independent Method. American Journal of Physical Anthropology 151, 290-301.
Harvati‐Papatheodorou, K. 2013. Neanderthals. A Companion to Paleoanthropology (ed. D. Begun), Wiley Blackwell, pp. 538-556.
Zaatari S., Harvati, K., Panagopoulou E. 2013. Occlusal molar microwear texture analysis and the diet of the Neanderthal from Lakonis. In S. Voutsaki and S. M. Valamoti (eds.): Diet, Economy and Society in the Ancient Greek World: Towards a Better Integration of Archaeology and Science. Pharos Supplement 1, Peters: Leuven, pp. 55-63.
Harvati K., Tourloukis V. 2013. Human Evolution in the Southern Balkans. Evolutionary Anthropology 22, 43-45.

Harvati K., Darlas A., Bailey S. E., Rein T. R.,El Zaatari S., Fiorenza L, Kullmer O., Psathi E. 2013. New Neanderthal remains from Mani peninsula, S. Greece: The Kalamakia Middle Palaeolithic cave site. Journal of Human Evolution
64, 486-499
Stoneking M., Harvati K. 2013. Early Old World migrations of Homo sapiens: human biology. Encyclopedia of Global Human Migration, Wiley Blackwell.
Rein T.R., Harvati K. 2013. Exploring third metacarpal capitate facet shape: implications for early hominin hand morphology. Anatomical Record 296, 240-249.
Harvati K. 2012. What happened to the Neanderthals? Nature Education Knowledge 3(10),13.
Freidline S.E., Gunz P., Harvati K., Hublin J.-J. 2012. Middle Pleistocene human facial morphology in an evolutionary and developmental context. Journal of Human Evolution 63, 723-740.
Harvati K. and Hublin J.-J. 2012. Morphological continuity of the face in the late Middle and Upper Pleistocene Hominins from North Western Africa – A 3-D geometric morphometric analysis. In: Hublin J.-J. and McPherron S. Modern Origins: A North African Perspective. Springer Verlag, pp. 179-188.
Frost, S.R., Schwartz, H.L., Giemsch, L., Morgan, L.E., Renne, P.R., Wildgoose M., Saanane C., Schrenk, F., Harvati K. 2012. Refined age estimates and paleoanthropological investigation oft he Manyara Beds, Tanzania. Journal of Anthropological Sciences 90,1-12.
Freidline S., Gunz P., Janković I., Harvati K., Hublin J.J. 2012. A comprehensive morphometric analysis of the frontal and zygomatic bone of the Zuttiyeh fossil from Israel. Journal of Human Evolution 62, 225–241
Singh N., Harvati K., Hublin J.-J., Klingenberg C. 2012. Morphological evolution through integration: A quantitative study of cranial integration in Homo, Pan, Gorilla and Pongo. Journal of Human Evolution 62:155-164.
Schwartz H., Renne P., Morgan L.E., Wildgoose M.M., Lippert P.C., Frost S.R., Harvati K., Schrenk F., Saanane C. 2012. Geochronology of the Manyara Beds, northern Tanzania: New tephrostratigraphy, magnetostratigraphy and 40Ar/39Ar ages. Quaternary Geochronology 7, 48-66.
Bastir M. , Rosas A., Gunz P., Pena-Melian A., Manzi G. , Harvati K., Kruszynski R., Stringer C., Hublin J.-J. 2011. Evolution of the base of the brain in highly encephalized human species. Nature Communications 2, 588
S. Benazzi, K. Douka, C. Fornai, C. C. Bauer, O. Kullmer, J. Svoboda, I. Pap, F.Mallegni, P. Bayle, M. Coquerelle, S. Condemi, A. Ronchitelli, K. Harvati & G. W.Weber. 2011. Early dispersal of modern humans in Europe and implications for Neanderthal behavior. Nature 479, 525-528
K. Harvati, C. Stringer, R. Grün, M. Aubert, P. Allsworth-Jones, C. Adebayo Folorunso. 2011. "Morphologie und Chronologie der Schädelkalotte aus Iwo Eleru (Nigeria, Later Stone Age)". PLoS ONE
Kranioti E., Holloway R., Ciprut T., Grigorescu D., Harvati K. 2011. A virtual assessment of the endocranial morphiology of the early modern European fossil calvaria from Cioclovina, Romania. Anatomical Record 294, 1083-1092 (Cover article)
Benazzi S., Viola B., Kullmer O., Fiorenza L., Harvati K., Metscher B, Paul T., Gruppioni G., Weber G., Mallegni F. 2011. A reassessment of the Neandertal teeth from Taddeo cave (southern Italy). Journal of Human Evolution 61, 377-387
Noback M., Harvati K., Spoor F. 2011. Climate related variation of the human nasal cavity. American Journal of Physical Anthropology 145, 599-614.
Gunz P. and Harvati K. 2011. Integration and homology of “chignon” and “hemibun” morphology. In: (Condemi S., ed.) 150 Years of Neanderthal Discoveries. Springer, p. 193-202.
Harvati K., Singh N. and Nicholson E. 2011. A 3-D view of the Neanderthal mandible. In: (Condemi S., ed.) 150 Years of Neanderthal Discoveries. Springer, p. 179-192.
Hubbe M., Harvati K., Neves W. 2011. Paleoamerican Morphology in the Context of European and East Asian Late Pleistocene Variation: Implications for Human Dispersion Into the New World. American Journal of Physical Anthropology 144, 442-453
Harvati K., Stringer C., Karkanas P. 2011. Multivariate analysis and classification of the Apidima 2 cranium from Mani, Southern Greece. Journal of Human Evolution 60, 246-250.
Harvati K. 2011. Homo heidelbergensis. McGraw-Hill 2011 Yearbook of Science & Technology. New York: McGraw-Hill, pp. 141-144.
Kaiser T., Seiffert C., Hertler C., Fiedler L., Schwartz H.L., Frost S.R., Giemsch L., Bernor R. L., Wolf D., Semprebon G., Nelson S.V., Schrenk F., Harvati K., Bromage T.G., Saanane C. 2010. Makuyuni, a new Lower Palaeolithic Hominid Site in Tanzania. Mitt. Hamb. Zool. Mus. Inst. 106, 69-110.
Allsworth-Jones P., Harvati K., Stringer C. 2010. The archaeological context of the Iwo Eleru cranium from Nigeria and preliminary results of new morphometric studies. BAR International Series 2164, 29-42.
Harvati K., Hublin J.-J. and Gunz P. 2010. Evolution of Middle-Late Pleistocene human cranio-facial form: A 3-D approach. Journal of Human Evolution 59, 445-464
Harvati, K. 2010 Neanderthals. Evo Edu Outreach 3, 367-376.
Hubbe M, Neves WA, Harvati K, 2010 Testing Evolutionary and Dispersion Scenarios for the Settlement of the New World. PLoS ONE 5(6): e11105.
Harvati K. 2009. Into Eurasia: Geometric morphometric analysis of the Upper Cave (Zhoukoudian) specimens. Journal of Human Evolution 57, 751-762.
Harvati K., Panagopoulou E., Runnels C. 2009. The Paleoanthropology of Greece. Evolutionary Anthropology 18, 131-143 (Cover article)
Hubbe M., Hanihara T., Harvati K. 2009. Climate signatures in the morphological differentiation of worldwide human populations. Anatomical Record 292, 1720-1733.
Harvati K. 2009. Petralona: Link between Africa and Europe? In: L. Schepartz, C. Bourbou, and S. Fox (Eds.). new Directions in the Skeletal Biology of Greece. Wiener Laboratory Series, ASCSA, Athens.
Nielsen-Marsh C., Hoffmann R., Toussaint M., Harvati K., Panagopoulou E., Hublin J-J., Richards M.P. 2009. Journal of Archaeological Science 36, 1758-1763.
Richards M., Grimes V., Smith C., Smith T., Harvati K., Hublin J.-J., Karkanas P., Panagopoulou E. 2009. Response to Nowell and Horstwood (2009) Journal of Archaeological Science 36, 1657-1658.
Smith T.M., Harvati K., Olejniczak A.J., Reid D.J., Hublin J.-J., Panagopoulou E. 2009. Dental development and enamel thickenss in the Lakonis Neanderthal molar. American Journal of Physical Anthropology 138, 112-118.
Richards M., Harvati K., Grimes V., Smith C., Smith T., Hublin J.-J., Karkanas P. and Panagopoulou E. 2008. Isotope evidence of Neanderthal mobility. Journal of Archaeological Science, 35, 1251-1256.
Harvati K., Panagopoulou E., Karkanas P., Athanassiou A. and Frost S. R. 2008. Preliminary results of the Aliakmon Paleolithic / Paleoanthropological Survey, Greece, 2004-2005. In Darlas A. and Mihailovic D., eds. The Paleolithic of the Balkans. BAR-IS (Oxford)
Harvati K., Gunz P. and Grigorescu D. 2007. Cioclovina (Romania): morphological affinities of an early modern European. Journal of Human Evolution.53, 732-746.
Tzedakis C., Hughen K, Cacho I. and Harvati K. 2007. Placing late Neanderthals in a climatic context. Nature, 449:206-208.
Harvati K. and Frost S.R. 2007. Dental eruption sequences in fossil colobines and the evolution of primate life histories. International Journal of Primatology, 28, 705-728.
Harvati K. 2007. Neanderthals and their contemporaries. In: Henke W. and Tattersall I. (eds.) Handbook of Paleoanthropology Vol. 3: Phylogeny of Hominines. Springer, pp. 1717-1748.
Grine F. E., Bailey R.M., Harvati K., Nathan R.P, Morris, A.G., Henderson G.M., Ribot I. and Pike A.W.G. 2007. Late Pleistocene Human Skull from Hofmeyr, South Africa and Modern Human Origins. Science 315, 226-229.
Gunz P. and Harvati K. 2007. The Neanderthal “chignon”: Variation, integration and homology. Journal of Human Evolution 52, 262-274.
Harvati K. 2007. 100 years of H. heidelbergensis - life and times of a controversial taxon. Mitteilungen der Gesellschaft für Urgeschichte 16, 85-94
Harvati K. 2007. The Krapina Neandertals: A Comprehensive, Centennial, Illustrated Bibliography. Journal of Human Evolution 52, 467.
Harvati K. and Weaver T. 2006. Reliability of cranial morphology in reconstructing Neanderthal phylogeny. In: K. Harvati and T. Harrison (eds.) Neanderthals Revisited: New Approaches and Perspectives. Springer, Dordrecht, pp. 239-254.
Harvati K. and Harrison T. 2006. Neanderthals Revisited. In: K. Harvati and T. Harrison (eds.) Neanderthals Revisited: New Approaches and Perspectives. Springer, Dordrecht, pp. 1-7.
Harvati K. and Weaver T. 2006. Human cranial anatomy and the differential preservation of population history and climate signatures. Anatomical Record 288A, 1225-1233.
Nicholson E. and Harvati K. 2006. Quantitative Analysis of Human Mandibular Shape Using 3-D Geometric Morphometrics. American Journal of Physical Anthropology 131, 368-383.
Panagopoulou E., Harvati K., Karkanas P., Athanassiou A., Elefanti P., Frost S.R. 2006. The West Macedonia Palaeolithic survey. The Archaeological Research in Macedonia and Thrace, 18, 631-640.
Delson E. and Harvati K. 2006. Return of the last Neanderthal. Nature. 443:762-763.
Panagopoulou, E., Karkanas, P., Kotjabopoulou, E., Harvati, K. and G. Tsartsidou. 2006 The Lakonis palaeolithic project. Archaeologicon Deltion, Chronika B'2(1999), 1003-1005.
Harvati K. , Frost S.R. and McNulty K.P. 2005. Neanderthal Variation and taxonomy – A reply to Ackermann (2005) and Ahern et al. (2005). Journal of Human Evolution 48: 653-660.
Harvati K. and Frost S. R. 2005. Dental eruption sequence in Mesopithecus pentelicus. Acta Palaeontologica Rom., vol. 5.
Reddy D. P., Harvati K. and Kim J. 2005. Alternative approaches to ridge-curve analysis using the example of the Neanderthal occipital ‘bun’. In: D. Slice (ed.) Modern Morphometrics in Physical Anthropology. Kluwer Academic Publishers.
Harvati K. 2005. Review of “Neanderthals and Modern Humans: An Ecological and Evolutionary Perspective” by Clive Finlayson. Human Biology.
Harvati K. , Frost S.R. and McNulty K.P. 2004. Neanderthal taxonomy reconsidered: Implications of 3D primate models of intra- and inter-specific differences. Proceedings of the National Academy of Sciences USA 101, 1147-1152.
Panagopoulou, E., Karkanas, P., Kotjabopoulou, E., Tsartsidou, Harvati, K., G. and Ntinou, M. 2004. Late pleistocene Archaeological and Fossil Human Evidence from Lakonis cave, Southern Greece. Journal of Field Archeology 29, 323-349.
Harvati K. 2004. 3-D geometric morphometric analysis of temporal bone landmarks in Neanderthals and modern humans. In: Elewa A. M. T. (ed.) Morphometrics, Applications in Biology and Paleontology. Springer-Verlag, p. 245-258.
Harvati K. , Panagopoulou E., Karkanas P. 2003. First Neanderthal remains from Greece: The evidence from Lakonis. Journal of Human Evolution 45, 465-473. DOI:10.1016/j.jhevol.2003.09.005
Harvati K. 2003 . Quantitative analysis of Neanderthal temporal bone morphology using 3-D geometric morphometrics. American Journal of Physical Anthropology 120, 323-338. DOI: 10.1002/ajpa.10122
Harvati K. 2003. The Neanderthal taxonomic position: models of intra- and inter-specific morphological variation. Journal of Human Evolution 44, 107-132. DOI:10.1016/S0047-2484(02)00208-7
Panagopoulou, E., Karkanas, P., Kotjabopoulou, E., Harvati, K. and G. Tsartsidou. 2003. The Lakonis palaeolithic project. Archaeologicon Deltion, Chronika (1997).
Harvati K. 2002. Models of shape variation between and within species and the Neanderthal taxonomic position: a 3D geometric morphometrics approach based on temporal bone morphology. British Archaeological Reviews International Series 1049, 25-30.
Delson E., Harvati K., Reddy D., Marcus L. F., Mowbray K., Sawyer G. J., Teuku Jacob and Márquez S. 2001. The Sambungmacan 3 Homo erectus calvaria: a comparative morphometric and morphological analysis. Anatomical Record 262, 380-397. DOI: 10.1002/ar.1048
Harvati K. 2000. Dental eruption sequence among colobine primates. American Journal of Physical Anthropology 112, 69-85.
Harvati K. 2000. Apidima. In: E. Delson, I. Tattersall, J. Van Couvering and A. S. Brooks (eds.) Encyclopedia of Human Evolution, 2nd edition, p.66-67. Garland Publishing, New York
Harvati K. and Delson E. 1999. Conference report: Paleoanthropology of the Mani peninsula. Journal of Human Evolution 36, 343-348.
Popular Science articles
In press
Harvati K. In press. Human Evolution in Greece. Proceedings of the conference: Myrtis: Face to Face with the Past, Akropolis Museum, Athens, May 2015 (in Greek).
Harvati K. In press. Neanderthals in the mirror: Contact, Extinction and Survival. Proceedings of the STARMUS conference, Tenerife, September 2014.
Harvati K. 2012. Der Geruchssinn. In: Von Sinnen: Wahrnehmung in der zeitgenössischen Kunst.Kunsthalle zu Kiel, Kerber Art, pp. 72-73.
Harvati K. 2009. Extinction and the Evolution of Humankind. In: What's Next: Dispatches on the Future of Science. Brockman, M. (ed.) Vintage Press.
Harvati K., Frost S.R. and McNulty K.P. 2006. Neandertaler: Vorfahren oder entfernte Verwandte? In: (Uelsberg, G., ed) Roots / Wurzeln der Menschheit, Reinisches Landesmuseum Bonn, pp. 133-139.

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